National Oceanic and
Atmospheric Administration
United States Department of Commerce



Current members (bold) and previous members (*) of the PMEL Ocean Molecular Ecology


Kelly, R.P., D.M. Lodge, K.N. Lee, S. Theroux, A.J. Sepulveda, C.A. Scholin, J.M. Craine, E. Andruszkiewicz Allan, K.M. Nichols, K.M. Parsons, K.D. Goodwin, Z. Gold, F.P. Chavez, R.T. Noble, C.L. Abbott, M.R. Baerwald, A.M. Naaum, P.M. Thielen, A.L. Simons, C.L. Jerde, J.J. Duda, M.E. Hunter, J.A. Hagan, R.S. Meyer, J.A. Steele, M.Y. Stoeckle, H.M. Bik, C.P. Meyer, E. Stein, K.E. James, A.C. Thomas, E. Demir-Hilton, M.A. Timmers, J.F. Griffith, M.J. Weise, and S.B. Weisberg (2023): Toward a national eDNA strategy for the United States. Environmental DNA, 10,

Gold, Z., A.O. Shelton, H.R. Casendino, J. Duprey, R. Gallego, A. Van Cise, M. Fisher, A.J. Jensen, E. D'Agnese, E. Andruszkiewicz Allan, A. Ramón-Laca, M. Garber-Yonts, M. Labare, K.M. Parsons, and R.P. Kelly (2023): Signal and noise in metabarcoding data. PLoS ONE, 18(5), e0285674,

Gold, Z., R.P. Kelly, A.O. Shelton, A.R. Thompson, K.D. Goodwin, R. Gallego, K.M. Parsons, L.R. Thompson, D. Kacev, and P.H. Barber (2023): Archived DNA reveals marine heatwave-associated shifts in fish assemblages. Environmental DNA, 00, 1-14,

Satterthwaite, E.V., A.E. Allen, R.H. Lampe, Z. Gold, A.R. Thompson, N. Bowlin, R. Swalethorp, K.D. Goodwin, E.L. Hazen, S.J. Bograd, S.A. Matthews, and B.X. Semmens (2023): Toward identifying the critical ecological habitat of larval fishes: An environmental DNA window into fisheries management. In: Frontiers in Ocean Observing: Emerging Technologies for Understanding and Managing a Changing Ocean. E.S. Kappel, V. Cullen, M.J. Costello, L. Galgani, C. Gordó-Vilaseca, A. Govindarajan, S. Kouhi, C. Lavin, L. McCartin, J.D. Müller, B. Pirenne, T. Tanhua, Q. Zhao, and S. Zhao, eds, Oceanography 36 (Supplement 1), 90–93,


Shelton, A.O., Z. Gold, E. D'Agnese, A. Van Cise, R. Gallego, K. Parsons, J. Duprey, H. Casedino, M. Garber-Yonts, M. Fisher, and R. Kelly (2022): Towards quantitative metabarcoding. Ecology, 104 (2), e3906,

Gold, Z., A.R. Wall, Teia M. Schweizer, N.D. Pentcheff, E.E. Curd, P.H. Barber, R.S. Meyer, R. Wayne, K. Stolzenbach, K. Prickett, J. Luedy, and R. Wetzer (2022): A manager's guide to using eDNA metabarcoding in marine ecosystems. PeerJ, 10, e14071,

Garber, A.I., C. R. Armbruster, S.E. Lee, V.S. Cooper, J.M. Bomberger, and S.M. McAllister (2022): SprayNPray: User-friendly taxonomic profiling of genome and metagenome contigs. MBC Genom., 23 (1), 1-8,


Galaska, M.P., G. Liu, D. West, K. Erickson, A. M. Quattrini, A. Bracco, and S. Herrera (2021): Seascape genomics reveals metapopulation connectivity network of Paramuricea biscaya in the northern Gulf of Mexico. Front. Mar. Sci., 8, 790929,

Galaska, M.P., D.S. Wethey, A. Arias, S.F. Dubois, K.M. Halanych, and S.A. Woodin (2021): The impact of aquaculture on the genetics and distribution of the onuphid annelid Diopatra biscayensis. Ecol. Evol., 11(11), 6184-6194,

Resh, C.A., M.P. Galaska, K.C. Benesh, J.P.A. Gardner, K.-J. Wei, R.-J. Yan, and A.R. Mahon (2021): Using genomics to link populations of an invasive species to its potential sources. Front. Ecol. Evol., 9, 575599,

Garber, A.I., G.A. Ramírez, S.M. McAllister, W. Orsi, and S. D’Hondt (2021): Cryptic metabolisms in anoxic subseafloor sediment. Environ. Microbiol. Rep., 13(5), 696-701,


Stepien, C.A.*, D. Leaman, and M.D. Niner. (2020): Chapter 4: Rhabdoviruses (with focus on Viral Hemorrhagic Septicemia Virus and other Novirhabdoviruses). In: Woo, P.T.K, J. Leong, & K. Buchmann. Climate Change and Infectious Fish Diseases. CABI, England, U.K.


Stepien, C.A.*, M.R. Snyder, A.E*. Elz. (2019): Invasion genetics of the silver carp Hypophthalmichthys molitrix across North America: Differentiation of fronts, introgression, and eDNA metabarcode detection. PLoS ONE, 14(3), e0203012.,

Marshall, N.T.*, C.A. Stepien*. (2019): Invasion genetics from eDNA and thousands of larvae: A targeted metabarcoding assay that distinguishes species and population variation of zebra and quagga mussels. Ecol. Evol., 9(6), 1-24,

Canonico, G., Buttigieg, P.L…Stepien, C.A.*, … et al. (2019): Global Observational Needs and Resources for Marine Biodiversity. Front. Mar. Sci., 6(367),

Marshall, N.*, and C.A. Stepien* (2019): Dreissenidae. Chapter 42 in Freshwater Mollusks of the World: A Distribution Atlas, C. Lydeard and K.S. Cummings (eds.), Johns Hopkins University Press.


Stepien, C.A.*, D.J. Eddins, M.R. Snyder*, and N.J. Marshall* (2018): Genetic change versus stasis over the time course of invasions: Trajectories of two concurrent, allopatric introductions of the Eurasian ruffe. Aquat. Invasions, 13(4), 537–552,

Stepien, C.A.*, M.R. Snyder*, and C.T. Knight (2018): Genetic divergence of nearby walleye spawning groups in central Lake Erie: Implications for management. N. Am. J. Fish. Manage., 38(4), 783–793,

Tajbakhsk, F., C.A. Stepien*, A. Abdoli, N. Tabatabaei, and B.H. Kiabi (2018): Geometric morphometric and meristic analysis of the deepwater goby, Ponticola bathybius (Kessler, 1877) (Teleostei: Gobiidae) in the Iranian waters of the Caspian Sea. Iran. J. Ichthyol., 5(1), 64–73.

Shepherd, B.S., A.R. Spear, A.M. Phillip, D.W. Leaman, C.A. Stepien*, O.J. Sepulveda-Villet, D.E. Palmquist, and M.M. Vijayan (2018): Effects of cortisol and lipopolysaccharide on expression of select growth-, stress- and immune-related genes in rainbow trout liver. Fish Shellfish Immunol., 74, 410-418,


Klymus, K.E., N.T. Marshall*, and C.A. Stepien* (2017): Environmental DNA (eDNA) metabarcoding assays to detect invasive invertebrate species in the Great Lakes. PLoS ONE, 12(5), e0177643,

Trebitz, A., J. Hoffman, J. Darling, E. Pilgrim, J. Kelly, E. Brown, W. Chadderton, S. Egan, E. Grey, S. Hashsham, K. Klymus, A. Mahon, J. Ram, M. Schjuiltz, C.A. Stepien*, and J. Schardt (2017): Early detection monitoring for aquatic non-indigenous species: Optimizing surveillance, incorporating advanced technologies, and identifying research needs. J. Environ. Manage., 202(1), 299–310,

Ke, Q., W. Weaver, A. Pore, B. Gorgoglione, J. Halo Wildschutte, P. Xiao, B.S. Shepherd, A. Spear, K. Malathi, C.A. Stepien*, V.N. Vakharia, and D.W. Leaman (2017): Role of Viral Hemorrhagic Septicemia Virus Matrix (M) protein in suppressing host transcription. J. Virol., 91(19), e00279-17,

Snyder, M.R.*, and C.A. Stepien* (2017): Genetic patterns across an invasion's history: A test of change versus stasis for the Eurasian round goby in North America. Mol. Ecol., 26(4), 1075–1090,

Stepien, C.A*., S.I. Karsiotis, T.J. Sullivan, and K.E. Klymus (2017): Population genetic structure and comparative diversity of smallmouth bass Micropterus dolomieu: Congruent patterns from two genomes. J. Fish Biol., 90(5), 2125–2147,